Two brothers ( who were priests ) received their portions for life, the shares of the other two were to revert to Owen on failure of issue.
2.
For surely the only harm to the individual animal from impairment of that " essential function " is not the failure of issue ( which harms only the issue ), but the psychic harm of perceiving that it will leave this world with no issue ( assuming, of course, that the animal in question, perhaps an endangered species of slug, is capable of such painful sentiments ).